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45. Brncic, T, Amarasekaran, B., McKenna, A., Mundry, R., Kühl, H.S. (2015) Large mammal diversity and their conservation in the human-dominated land-use mosaic of Sierra Leone. /Biodiversity and Conservation/. DOI 10.1007/s10531-015-0931-7 [pdf]

44. Heinicke, S., Kalan, A. K., Wagner, O. J., Mundry, R., Lukashevich, H., Kühl, H. S. (2015). Assessing the performance of a semi‐automated acoustic monitoring system for primates. /Methods in Ecology and Evolution/. DOI: 10.1111/2041-210X.12384 [pdf]

43. Kalan, A. K., Mundry, R., Wagner, O. J., Heinicke, S., Boesch, C., Kühl, H. S. (2015). Towards the automated detection and occupancy estimation of primates using passive acoustic monitoring. /Ecological Indicators/, /54/, 217-226. [pdf]

42. Saéz, A.M., Weiss, S., Nowak, K., Lapeyre, V., Zimmermann, F., Düx, A., Kühl, H.S., Kaba, M., Regnaut, S., Merkel, K., Sachse, A., Thiesen, U., Villányi, L., Boesch, C., Dabrowski, P.W., Radonić, A., Nitsche, A., Leendertz, S.A.J., Petterson, S., Becker, S., Krähling, V., Couacy-Hymann, E., Akoua-Koffi, C., Weber, N., Schaade, L., Fahr, J., Borchert, M., Gogarten, J. F., Calvignac-Spencer, S., Leendertz, F.H. (2015) Investigating the zoonotic origin of the West African Ebola epidemic. EMBO Molecular Medicine. doi: 10.15252/emmm.201404792. [pdf]

41. Junker, J., Boesch, C., Mundry, R., Stephens, C., Lormie, M., Tweh, C., Kühl, H. S. (2015). Education and access to fish but not economic development predict chimpanzee and mammal occurrence in West Africa. Biological Conservation, 182, 27-35. [pdf]


40. Tranquilli, S., Abedi-Lartey, M., Abernethy, K., Amsini, F., Asamoah, A., Balangtaa, C., Blake, S, Bouanga, E., Breuer, T., Brncic, T.M., Campbell, G., Chancellor, R., Chapman, C.A., Davenport, T. R. B. Dunn, A., Dupain, J., Ekobo, A., Eno-Nku, M, Etoga, G., Furuichi, F., Gatti, S., Ghiurghi, A., Hashimoto, C., Hart, J.A., Head, J., Hega, M, Herbinger, I, Hicks, T.C., Holbech, L.H., Huijbregts, B., Kühl, H.S., Imong, I., Le-Duc Yeno, S., Linder, J., Marshall, P., Lero, P.M., Morgan, D., Mubalama, L., N’Goran, P.K., Nicholas, A., Nixon, S., Normand, E., Nziguyimpa, L., Nzooh-Dongmo, Z., Ofori-Amanfo, R., Ogunjemite, B.G., Petre, C.-A., Rainey, H.J., Regnaut, S., Robinson, O., Rundus, A., Sanz, C.M., Okon, D.T., Todd, A., Warren, I., Sommer, V. (2014) Protected Areas in Tropical Africa: Assessing Threats and Conservation Activities. PLoS ONE, 9, e114154. [pdf]

39. Roy, J., Vigilant, L., Gray, M., Wright, E., Kato, R., Kabano, P., Basabose, A., Tibenda, E., Kühl, H.S., Robbins, M. M. (2014). Challenges in the use of genetic mark-recapture to estimate the population size of Bwindi mountain gorillas (Gorilla beringei beringei). Biological Conservation, 180, 249-261. [pdf]

38. Wich, S. A., Garcia-Ulloa, J., Kühl, H. S., Humle, T., Lee, J. S., Koh, L. P. (2014). Will Oil Palm’s Homecoming Spell Doom for Africa’s Great Apes? Current Biology, 24, 1659-1663. [pdf]

37. Imong, I., Robbins, M.M., Mundry, R., Bergl, R., Kühl, H.S. (2014) Informing conservation management about structural vs. functional connectivity: a case study of Cross River gorillas. American Journal of Primatology, 76, 978-988. [pdf]

36. Serckx, A., Huynen, M. C., Bastin, J. F., Hambuckers, A., Beudels-Jamar, R. C., Vimond, M., Raynaud, E., Kühl, H. S. (2014). Nest Grouping Patterns of Bonobos (Pan paniscus) in Relation to Fruit Availability in a Forest-Savannah Mosaic. PLOS ONE, 9, e93742. [pdf]

35. Neba Funwi-Gabga, Hjalmar S. Kuehl, Fiona G. Maisels, Susan M. Cheyne, Serge A. Wich and Elizabeth A. WilliamsonThe status of apes across Africa and Asia Rainer, H., White, A., & Lanjouw, A. (Eds.). (2014). Extractive Industries and Ape Conservation. Cambridge University Press. [pdf]

34. Tweh, C.G., Lormie, M.M., Kouakou, C.Y., Hillers, A., Kühl, H.S., Junker (2014) Conservation status of chimpanzees (Pan troglodytes verus) and other large mammals in Liberia: a nationwide survey. Oryx DOI: [pdf]

33. Fünfstück, T., Arandjelovic, M., Morgan, D. B., Sanz, C., Breuer, T., Stokes, E. J., Patricia Reed, Sarah H. Olson, Ken Cameron, Alain Ondzie, Martine Peeters, Hjalmar S. Kühl, Cipolletta, C., Todd, A., Masi, S., Doran-Sheehy, D.M., Bradley, B.J., Vigilant, L. (2014). The genetic population structure of wild western lowland gorillas (Gorilla gorilla gorilla) living in continuous rain forest. American Journal of Primatology. DOI: 10.1002/ajp.22274. [pdf]

32. Imong, I., Robbins, M. M., Mundry, R., Bergl, R., Kühl, H. S. (2014). Distinguishing ecological constraints from human activity in species range fragmentation: the case of Cross River gorillas. Animal Conservation.  DOI: 10.1111/acv.12100 [pdf]

31. Hicks, T. C., Tranquilli, S., Kuehl, H., Campbell, G., Swinkels, J., Darby, L., Boesch, C., Hart, J., Menken, S. B. (2014). Absence of evidence is not evidence of absence: Discovery of a large, continuous population of Pan troglodytes schweinfurthii in the Central Uele region of northern DRC. Biological Conservation, 171, 107-113. [pdf]


30. Hickey, J.R., Nackoney, J., Nibbelink, N.P., Blake, S.,Bonyenge, A., Coxe, S., Dupain, J., Emetshu. M., Furuichi, T., Grossmann, F., Guislain, P., Hart, J., Hashimoto, C., Ikembelo, B., Ilambu, O., Inogwabini, B.I., Liengola., I., Lokasola, A.L., Lushimba, A., Maisels, F., Masselink, J., Mbenzo, V., Mulavwa, N.M., Naky, P., Ndunda, N.M., Nkumu, P., Omasombo, V., Reinartz, G.E., Rose, R., Sakamaki, T., Strindberg, S., Takemoto, H., Vosper, A., Kühl, H.S. (2013) Human proximity and habitat fragmentation are key drivers of the rangewide bonobo distribution. Biodiversity and Conservation 22, 3085-3104. [pdf]

29. Murai, M., Ruffler,H., Berlemont, A., Campbell, G., Esono, F., Agbor, A., Mbomio, D., Ebana, A., Nze, A., Kühl, H.S. (2013) Priority areas for large mammal conservation in Equatorial Guinea. PLoS ONE 8, e75024 [pdf]

28. Kühl, H.S., Burghardt, T. (2013) Fractal representation and recognition for animal biometrics: a reply to Jovani et al. Trends in Ecology and Evolution 28, 500-501. [pdf]

27. Head, J.S., Boesch, C., Robbins, M.M., Rabanal, L.I., Makaga, L., Kühl, H.S. (2013) Effective socio-demographic population assessment of elusive species for ecology and conservation management. Ecology and Evolution. doi: 10.1002/ece3.670 [pdf]

26. N’Goran, P.K., Kouakou, C. Y., N’goran, E.K., Konaté, S., Herbinger, I., Yapi, F.A., Kuehl, H., Boesch, C. (2013) Chimpanzee conservation status in the World Heritage Site Taï National Park, Côte d’Ivoire.International Journal of Innovation and Applied Studies, 3, 326-336. [pdf]

25. Kühl, H.S., Burghardt, T. (2013) Animal biometrics: quantifying and detecting phenotypic appearance. Trends in Ecology and Evolution 28, 432–441 [pdf]

24. Calvignac-Spencer, S., Merkel, K., Kutzner, N., Kühl, H., Boesch, C., Kappeler, P.M., Metzger, S., Schubert, G., Leendertz, F. (2013) Carrion fly-derived DNA as a tool for comprehensive and cost-effective assessment of mammalian biodiversity. Molecular Ecology 22, 915–924. [pdf]


23. Junker, J., Blake, S., Boesch, C., Campbell, G., du Toit, L., Duvall, C., Ekobo, A., Etoga, G., Galat-Luong, A., Gamys, J., Ganas-Swaray, J., Gatti, S., Ghiurghi, A., Granier, N., Hart, J., Head, J., Herbinger, I., Hicks, T.C., Huijbregts, B., Imong, I.S., Kuempel, N., Lahm, S., Lindsell, J., Maisels, F., McLennan, M., Martinez, L., Morgan, B., Morgan, D.,Mulindahabi, F., Mundry, R., N’Goran, P.K., Normand, E., Ntongho, A., Okon, D.T., Petre, C.A., Plumptre, A., Rainey, H., Regnaut, S., Sanz, C., Stokes, E., Tondossama, A.,Tranquilli, S., Sunderland-Groves, J., Walsh, P., Warren, Y., Williamson, E.A., Kuehl, H.S. (2012) Recent decline in suitable environmental conditions for African great apes. Diversity and Distribution. DOI: 10.1111/ddi.12005 [pdf] [Supplementary Material Part 1] [Supplementary Material Part 2]

22. Kühl, H.S., N'Guessan, A., Riedel, J., Metzger, S., Deschner, T. (2012) The Effect of Climate Fluctuation on Chimpanzee Birth Sex Ratio. PLoS ONE 7(4): e35610. doi:10.1371/journal.pone.0035610 [pdf]

21. N'Goran, P.K., Boesch, C., Mundry, R., N'Goran, E. K., Herbinger, I., Yapi, F.A., Kühl, H.S. (2012) Hunting, law enforcement and African primate conservation. Conservation Biology. DOI: 10.1111/j.1523-1739.2012.01821. [pdf] [Supplementary Material Part 1] [Supplementary Material Part 2]

20. Wich, S.A., Fredriksson, G.M., Usher, G., Peters, H.H., Priatna, D., Basalamah, F., Susanto, W., Kühl, H.S. (2012) Hunting of Sumatran orang-utans and its importance in determining distribution and density. Biological Conservation 146, 163-169. [pdf] [Supplementary Material]

19. Tranquilli, S., Abedi-Lartey, M., Amsini, F., Arranz, L., Asamoah, A., Babafemi, O., Barakabuye, N., Campbell, G., Chancellor, R., Davenport, T.R.B., Dunn, A., Dupain, J., Ellis, C., Etoga, G., Furuichi, T., Gatti, S., Ghiurghi, A., Greengrass, E., Hashimoto, C., Hart, J., Herbinger, I., Hicks, T.C., Holbech, L.H., Huijbregts, B., Imong, I., Kumpel, N., Maisels, F., Marshall, P., Nixon, S., Normand, E., Nziguyimpa, L., Nzooh-Dogmo, Z., Tiku Okon, D., Plumptre, A., Rundus, A. Sunderland-Groves, J., Todd, A., Warren, Y., Mundry, R., Boesch, C., Kuehl, H.S. (2012) Lack of conservation effort rapidly increases African great ape extinction risk. Conservation Letters 5, 48-55. [pdf] [Supplementary Material]


18. Kouakou, C.Y., Boesch, C., Kuehl, H.S. (2011) Identifying hotspots of chimpanzee group activity from transect surveys in Taï National Park, Côte d'Ivoire. Journal of Tropical Ecology 27: 621-630. [pdf]

17. Hoppe-Dominik, B., Kühl, H.S., Radl, G., Fischer, F. (2011) Long-term monitoring of large rainforest mammals in the Biosphere Reserve of Taï National Park, Côte d’Ivoire. African Journal of Ecology doi: 10.1111/j.1365-2028.2011.01277.x [pdf]

16. Campbell, G., Kuehl, H., Diarrassouba, A., N'Goran, P. K. and C. Boesch (2011) Long-term research sites as refugia for threatened and over-harvested species. Biology Letters doi: 10.1098/rsbl.2011.0155 [pdf] [Supplementary Material]


15. Spehar, N.S., Mathewson, P.D., Nuzuar, Wich, S.A., Marshall, A.J., Kühl, H., Nardiyono, Meijaard, E. (2010) Estimating Orangutan densities using the standing crop and marked nest count methods: Lessons learned for conservation. Biotropica 1-10. [pdf]

14. Arandjelovic M., Head, J.., Kuehl, H., Boesch, C., Robbins, M.M., Maisels, F., Vigilant, L. Effective non-invasive genetic monitoring of multiple wild western gorilla groups. Biological Conservation 143: 1780–1791. [pdf]

13. Rabanal, L., Kuehl, H.S. Mundry, R., Robbins, M., Boesch, C. (2010) Oil prospecting and its impact on large rainforest mammals in Loango National Park, Gabon. Biological Conservation. 143: 1017-1024. [pdf]


12. Junglen S., Kurth A., Ellerbrok H., Pauli G., Nunn C., Kuehl H., Leendertz F. H. (2009) Examining landscape factors influencing relative distribution of mosquito genera and frequency of virus infection. EcoHealth 6: 239-249. [pdf]

11. Kouakou, Y.C., Boesch, C., Kuehl, H. S.: (2009) Estimating chimpanzee population size with nest counts: Validating methods in Taï National Park, American Journal of Primatology 71: 447-457. [pdf]

10. Kuehl, H. S.; Nzeingui, C.; Le Duc Yeno, S.; Huijbregts, B.; Boesch, C.; Walsh, P. D. (2009): Discriminating between village and commercial hunting of apes. Biological Conservation 142: 1500-1506. [pdf]


9. Campbell, G., Kuehl, H.S., Ngoran, P, Boesch, C. (2008) Alarming decline of West African chimpanzees in Côte d’Ivoire. Current Biology 18: 904-905. [pdf]

8. Kuehl, H.S, Maisels, F., Williamson, L., Ancrenaz, M. (2008) Best Practice Guidelines for Surveys and Monitoring of Great Ape Populations. Occasional papers of the IUCN Species Survival Commission. [pdf in English | pdf in French]

7. Todd, A.F., Kuehl, H.S, Cipolletta, C., Walsh, P.D. (2008) Using dung to estimate gorilla density: Modeling dung production rate. International Journal of Primatology 29: 549-563. [pdf]

6. Kuehl, H.S., Elzner, C., Moebius, Y., Boesch, C., Walsh, P.D. (2008) The price of play: self-organized infant mortality cycles in chimpanzees. PloS ONE 3: e2440 [pdf]

5. Köndgen, S., Kühl, H.S., Ngoran, P., Walsh, P.D., Schenk, S., Ernst, N., Biek, R., Formenty, P., Mätz-Rensing, K., Schweiger, B., Junglen, S., Ellerbrok, H., Nitsche, A.,Briese, T.,Lipkin, WI., Pauli, G.,Boesch, C., Leendertz FH (2008) Pandemic human viruses cause decline of endangered great apes. Current Biology 18: 260-264. [pdf | Supplemental data]


4. Kuehl, H.S, Todd, A., Boesch, C., and Walsh, P.D. (2007) Manipulating dung decay time for efficient large-mammal density estimation: Gorillas and dung height. Ecological Applications 17: 2403–2414. [pdf]

3. Kuehl, H.S, Williamson, L, Sanz, C, Morgan, D., Boesch, C. (2007) Launch of A.P.E.S. database. Gorilla Journal 34: 20-21. [pdf]


2. Wimmer, R., Kühl, H., Röttger. S., Schempp, W. (2002) Comparative mapping of CDY and DAZ in higher primates. Cytogenetic and Genome Research 96: 287–289 [pdf]


1. Kühl, H., Röttger, S., Heilbronner, H., Enders, H, Schempp, W. (2001) Loss of the Y chromosomal PAR2-region in four familial cases of satellited Y chromosomes (Yqs). Chromosome Research 9: 215-222. [pdf]



Vaidyanathan, G. (2011) The cultured chimpanzees. Do chimpanzees have traditions? As wild populations dwindle, researchers are racing to find out. Nature 476: 266-269. [pdf]