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Mark Stoneking

Max Planck Institute for Evolutionary Anthropology
Department of Evolutionary Genetics
Deutscher Platz 6
D-04103 Leipzig

Tel: +49 (0)341 3550 502
Fax: +49 (0)341 3550 555

E-mail: stonekingeva.mpgde
Website: http://www.eva.mpg.de/genetics/

Curriculum Vitae
Vice Ombudsperson

Curriculum Vitae

date of birth: August 1, 1956

Download CV Mark Stoneking

Educational History

University of Oregon1974-1977AnthropologyB.A.
The Pennsylvania State University1977-1979GeneticsM.S.
University of Wisconsin, Madison1979-1981Genetics-
University of California, Berkeley1981-1986GeneticsPh.D.

Professional History

Group Leader, Max Planck Institute for Evolutionary Anthropology, and Honorary Professor of Biological Anthropology, University of Leipzig, Leipzig, Germany
1998Professor of Anthropology, Pennsylvania State University
1996-1997Visiting Professor, Zoology Institute, University of Munich
1994-1998Associate Professor of Anthropology, Pennsylvania State University
1990-1994 Assistant Professor, Dept. of Anthropology, Graduate Program in Genetics,
and Institute of Molecular Evolutionary Genetics, Pennsylvania State University
1989-1990 Associate Research Scientist, Dept. of Human Genetics,
Cetus Corporation, Emeryville, California
1989Staff Scientist, Human Genome Center,
Lawrence Berkeley Laboratory, Berkeley
1986-1988Postdoctoral Fellow, Dept. of Biochemistry,
University of California, Berkeley

Fellowships, Prizes and Awards

National Science Foundation Graduate Fellowship, 1977-1978, 1979-1981
Pennsylvania State University Graduate Fellowship, 1978-1979
Ernest Brown Babcock Scholarship, University of California, Berkeley, 1985-1986
John Belling Prize in Genetics, University of California, Berkeley, 1990
University of Oregon Outstanding Young Alumnus Award, 1990
Japanese Society for the Promotion of Science Fellowship, 1995
FBI Award for Service to the Forensic DNA Community, 1998
Fellow, American Association for the Advancement of Science, 2000
Visiting Professor, CNRS Laboratory for Biometry and Evolutionary Biology, Lyon, France, February 2016
Eugène Dubois Chair, University of Maastricht, Maastricht, The Netherlands, 2016
Elected to the National Academy of Sciences USA, 2020


Eastern Indonesia, January-February 1988
Solomon Islands, August-September 2004
Mindanao, Philippines, August 2005
Zambia, August-September 2007
Namibia, May-June 2011
Botswana and Namibia, July-August 2015 (returning genetic results to participating communities)

Professional Activities

Associate Editor: Journal of Human Evolution, 1990-1993; Human Biology, 1993-1997; BioEssays, 2004-2013; Anthropological Science, 2004-2013; Evolutionary Biology, 2007-present; BMC Genetics, 2008-2015; Gene, 2009-2010; Investigative Genetics, 2009-2016; EMBO Reports, 2010-present; Language Dynamics and Change, 2010-present.

Senior Editor, Annals of Human Genetics, 2008-2013.

Technical Working Group, DNA Analysis Methods, FBI, 1993-1998

Defense Science Board Task Force on DNA Technology for Identification of Ancient Remains, 1994-1995

Wellcome Trust Bioarchaeology Panel, 2001-2005

Steering Committee for National Energy Research Council (NERC) Program on Environmental Factors and the Chronology of Human Evolution and Dispersal (EFCHED), 2001-2006

Review Panel, Wenner-Gren Foundation for Anthropological Research, 2007-2008

Advisory Committee, The Role of Culture in Early Expansions of Humans, Heidelberg Academy of Sciences, 2008-present

Advisory Board, US National Evolutionary Synthesis Center, 2011-2014

Chair, Scientific Advisory Committee of the Program on Forensics and Ethnicity, Philippine Genome Center, 2011-present

Review Panel, Portuguese Foundation for Science and Technology, 2013-2014

Scientific Advisory Board, IDEXLYON Program, 2017-present


This publication list is currently beeing updated. The process will be finished in a few minutes.


Stoneking, M. and H. Soodyall. 1996. Human evolution and the mitochondrial genome. Current Opinion in Genetics and Development 6:731-736.

Erlich, H. A., T. F. Bergström, M. Stoneking, and U. Gyllensten. 1996. HLA sequence polymorphism and the origin of humans. Science 274:1552-1554.

Thomas, R., H. Zischler, S. Pääbo, and M. Stoneking. 1996. Novel mitochondrial DNA insertion polymorphism and its usefulness for human population studies. Human Biology 68:847-854. 

Stone, A. C. and M. Stoneking. 1996. Genetic analyses of an 8000 year-old native American skeleton. Ancient Biomolecules 1:83-87.

Soodyall, H., T. Jenkins, R. Hewitt, A. Krause, and M. Stoneking. 1996. The peopling of Madagascar. In: Molecular Biology and Human Diversity, A. J. Boyce and C. G. N. Mascie-Taylor (eds.), Cambridge University Press, Cambridge, pp.156-170.

Melton, T. and M. Stoneking. 1996. Extent of heterogeneity in mitochondrial DNA of ethnic Asian populations. Journal of Forensic Sciences 41:591-602.

Stoneking, M. 1996. Mitochondrial DNA heteroplasmy: out of the closet. Biol. Chem. 377:603-604.

Stoneking, M. 1996. Mitochondrial DNA variation and human evolution. In: Human Genome Evolution, M. Jackson, T. Strachan, and G. Dover (eds.), BIOS Scientific Publishers, Oxford, pp. 263-281.

Bonatto, S. L., A. J. Redd, F. M. Salzano, and M. Stoneking. 1996. Lack of ancient Polynesian-Amerindian contact. American Journal of Human Genetics 59:253-255.

Knight, A., M. A. Batzer, M. Stoneking, H. K. Tiwari, W. D. Scheer, R. J. Herrera, and P. L. Deininger. 1996. DNA sequences of Alu elements indicate a recent replacement of the human autosomal genetic complement. Proceedings of the National Academy of Sciences USA 93:4360-4364.

Soodyall, H., L. Vigilant, A. V. Hill, M. Stoneking, and T. Jenkins. 1996. Mitochondrial DNA control region sequence variation suggests multiple independent origins of an "Asian-specific" 9-bp deletion in sub-Saharan Africans. American Journal of Human Genetics 58:595-608.

Nachman, M. W., W. M. Brown, M. Stoneking, and C. F. Aquadro. 1996. Non-neutral mitochondrial DNA variation in humans and chimpanzees. Genetics 142:953-963.

Batzer, M. A., S. S. Arcot, J. W. Phinney, M. Alegria-Hartman, D. H. Kass, S. M. Milligan, C. Kimpton, P. Gill, M. Hochmeister, P. A. Ioannou, R. J. Herrera, D. A. Boudreau, W. D. Scheer, B J. B. Keats, P. L. Deininger, and M. Stoneking. 1996. Genetic variation of recent Alu insertions in human populations. Journal of Molecular Evolution 42:22-29.

Bonnichsen, R., M. T. Beatty, M. D. Turner, and M. Stoneking. 1996. What can be learned from hair? A hair record from the Mammoth Meadow locus, southwestern Montana. In: Prehistoric Mongoloid Dispersals, T. Akazawa and E. J. E. Szathmary (eds.), Oxford University Press, Oxford, pp. 201-213.

Stone, A. C., G. R. Milner, S. Pääbo, and M. Stoneking. 1996. Sex determination of ancient human skeletons using DNA. American Journal of Physical Anthropology 99:231-238.


Novick, G. E., C. C. Novick, J. Yunis, E. Yunis, K. Martinez, G. G. Duncan, G. M. Troup, P. L. Deininger, M. Stoneking, M. A. Batzer, and R. J. Herrera. 1995. Polymorphic human specific Alu insertions as markers for human identification. Electrophoresis 16:1596-1601.

Stoneking, M. 1995. Ancient DNA: How do you know when you have it and what can you do with it? American Journal of Human Genetics 57:1259-1262.

Soodyall, H., M. Stoneking, and T. Jenkins. 1995. ‘Polynesian’ mtDNA in the Malagasy. Nature Genetics 10:377-378.

Melton, T., R. Peterson, A. Redd, N. Saha, A. S. M. Sofro, J. Martinson, and M. Stoneking. 1995. Polynesian genetic affinities with southeast Asian populations as identified by mtDNA analysis. American Journal of Human Genetics 57:403-414.

Redd, A. J., N. Takezaki, S. T. Sherry, S. T. McGarvey, A. S. M. Sofro, and M. Stoneking. 1995. Evolutionary history of the COII/tRNAlys intergenic 9 b.p. deletion in human mitochondrial DNAs from the Pacific. Molecular Biology and Evolution 12:604-615.

Gill, P., C. Kimpton, R. Aliston-Greiner, K. Sullivan, M. Stoneking, T. Melton, J. Nott, S. Barritt, R. Roby, M. Holland, and V. Weedn. 1995. Establishing the identity of Anna Anderson Manahan. Nature Genetics 9:9-10.


Connor, A. and M. Stoneking. 1994. Assessing ethnicity from human mitochondrial DNA types determined by hybridization with sequence-specific oligonucleotides. Journal of Forensic Sciences 39:1360-1371.

Batzer, M. A., M. Stoneking, M. Alegria-Hartman, H. Bazan, D. H. Kass, T. H. Shaikh, G. E. Novick, P. A. Ionnou, W. D. Scheer, R. J. Herrera, and P. L. Deininger. 1994. African origin of human-specific polymorphic Alu insertions. Proceedings of the National Academy of Sciences USA 91: 12288-12292.

Weiss, K. M., A. V. Buchanan, C. Daniel, and M. Stoneking. 1994. Optimizing utilization of DNA from rare or archival anthropological samples. Human Biology 66: 789-804.

Sherry, S. T., A. R. Rogers, H. Harpending, H. Soodyall, T. Jenkins, and M. Stoneking. 1994. Mismatch distributions of mtDNA reveal recent human population expansions. Human Biology 66: 761-775.

Cheng, S., R. Higuchi, and M. Stoneking. 1994. Complete mitochondrial genome amplification. Nature Genetics 7: 350-351.

Stoneking, M. 1994. In defense of “Eve” - a response to Templeton's critique. American Anthropologist 96: 131-141.

Stoneking, M. 1994. Mitochondrial DNA and human evolution.  Journal of Bioenergetics and Biomembranes 26: 241-249.


Batzer, M. A., M. Alegria-Hartman, H. Bazan, D. H. Kass, T. H. Shaikh, G. E. Novick, P. A. Ioannou, D. A. Boudreau, W. D. Scheer, R. J. Herrera, M. Stoneking, and P. L. Deininger. 1993. Alu repeats as markers for human population genetics. In: Proceedings of the Fourth International Symposium on Human Identification, Promega Publishing, Madison, Wisconsin, pp. 49-57.

Stone, A. C. and M. Stoneking. 1993. Ancient DNA from a pre-Columbian Amerindian population. American Journal of Physical Anthropology 92: 463-471.

Wilson, M. R., M. Stoneking, M. M. Holland, J. A. DiZinno, and B. Budowle. 1993. Guidelines for the use of mitochondrial DNA sequencing in forensic science. Crime Laboratory Digest 20: 68-77.

Hutchin, T., I. Haworth, K. Higashi, N. Fischel-Ghodsian, M. Stoneking, N. Saha, K. Arnos, and G. Cortopassi. 1993. A molecular basis for human hypersensitivity to aminoglycoside antibiotics. Nucleic Acids Research 21: 4174-4179.

Stoneking, M. 1993. DNA and recent human evolution. Evolutionary Anthropology 2: 60-73.

Harpending, H. C., S. T. Sherry, A. R. Rogers, and M. Stoneking. 1993. The genetic structure of ancient human populations. Current Anthropology 34: 483-496.

Buchanan, A. V., S. T. Sherry, K. M. Weiss, S. T. McGarvey, J. V. Neel, and M. Stoneking. 1993. Extraction of DNA from frozen red blood cells. Human Biology 65: 647-654.


Stoneking, M., S. T. Sherry, and L. Vigilant. 1992. Geographic origin of human mitochondrial DNA revisited. Systematic Biology 41: 384-391.

Moonsamy, P. V., V. C. Suraj, T. L. Bugawan, R. K. Saiki, M. Stoneking, J. Roudier, M. M. A. Magzoub, A. V. S. Hill, and A. B. Begovich. 1992. Genetic diversity within the HLA Class II region: ten new DPB1 alleles and their population distribution. Tissue Antigens 40: 153-157.

Stoneking, M., S. T. Sherry, A. J. Redd, and L. Vigilant. 1992. New approaches to dating suggest a recent age for the human mtDNA ancestor. Phil. Trans. R. Soc. Lond. B 337: 167-175.

Perna, N. T., M. A. Batzer, P. L. Deininger, and M. Stoneking. 1992. Alu insertion polymorphism: a new type of marker for human population studies. Human Biology 64: 641-648. 

Hedges, S. B., S. Kumar, K. Tamura, and M. Stoneking. 1992. Human origins and analysis of mitochondrial DNA sequences. Science 255: 737-739.


Wilson, A.C., M. Stoneking, and R. L. Cann. 1991. Ancestral geographic states and the peril of parsimony. Systematic Zoology 40: 363-365.

Vigilant, L., M. Stoneking, H. Harpending, K. Hawkes, and A. C. Wilson. 1991. African populations and the evolution of human mitochondrial DNA. Science 253: 1503-1507.

Stoneking, M., D. Hedgecock, R. G. Higuchi, L. Vigilant, and H. A. Erlich. 1991. Population variation of human mitochondrial DNA control region sequences detected by enzymatic amplification and sequence-specific oligonucleotide probes. American Journal of Human Genetics 48: 370-382.


Stoneking, M. 1990. Departure of human mtDNA variation from neutral expectations: an alternative explanation. Journal of Molecular Evolution 31: 343-346.

Helmuth, R., N. Fildes, E. Blake, M.C. Luce, J. Chimera, R. Madej, C. Gorodezky, M. Stoneking, N. Schmill, W. Klitz, R. Higuchi, and H. A. Erlich. 1990. HLA-DQ allele and genotype frequencies in various human populations, determined by using enzymatic amplification and oligonucleotide probes. American Journal of Human Genetics 47: 515-523.

Stoneking, M., L.B. Jorde, K. Bhatia, and A.C. Wilson. 1990. Geographic variation in human mitochondrial DNA from Papua New Guinea. Genetics 124: 717-733.


Stoneking, M. and A.C. Wilson. 1989. Mitochondrial DNA. In: The Colonization of the Pacific: A Genetic Trail, A.V.S. Hill and S.W. Serjeantson (eds.), Oxford University Press, Oxford, pp. 215-245.

Stoneking, M. and R.L. Cann. 1989. African origin of human mitochondrial DNA. In: The Human Revolution: Behavioural and Biological Perspectives on the Origins of Modern Humans, P. Mellars and C. Stringer (eds.), Edinburgh University Press, Edinburgh, pp. 17-30.


Vigilant, L., M. Stoneking, and A.C. Wilson. 1988. Conformational mutation in human mtDNA detected by direct sequencing of enzymatically amplified DNA. Nucleic Acids Research 16: 5945-5955.


Wilson, A.C., M. Stoneking, R.L. Cann, E.M. Prager, S.D. Ferris, L.A. Wrischnik, and R.G. Higuchi. 1987. Mitochondrial clans and the age of our common mother. In: Human Genetics, Proceedings of the Seventh International Congress, Berlin 1986, F. Vogel and K. Sperling (eds.), Springer-Verlag, Berlin, pp. 158-164.

Wrischnik, L.A., R.G. Higuchi, M. Stoneking, H.A. Erlich, N. Arnheim, and A.C. Wilson. 1987. Length mutations in human mitochondrial DNA: direct sequencing of enzymatically amplified DNA. Nucleic Acids Research 15: 529-542.

Cann, R.L., M. Stoneking, and A.C. Wilson. 1987. Disputed African origin of human populations. Nature 329: 111-112.

Cann, R.L., M. Stoneking, and A.C. Wilson. 1987. Mitochondrial DNA and human evolution. Nature 325: 31-36.


Whittam, T.S., A.G. Clark, M. Stoneking, R.L. Cann, and A.C. Wilson. 1986. Allelic variation in human mitochondrial genes based on patterns of restriction site polymorphism. Proceedings of the National Academy of Science USA 83: 9611-9615.

Stoneking, M., K. Bhatia, and A.C. Wilson. 1986. Rate of sequence divergence estimated from restriction maps of mitochondrial DNAs from Papua New Guinea. Cold Spring Harbor Symposia on Quantitative Biology 51: 433-439.

Stoneking, M., K. Bhatia, and A.C. Wilson. 1986. Mitochondrial DNA variation in Eastern Highlanders of Papua New Guinea. In: Genetic Variation and its Maintenance, D.F. Roberts and G. DeStefano (eds.), Cambridge University Press, Cambridge, pp. 87-100.


Wilson, A.C., R.L. Cann, S.M. Carr, M. George, U.B. Gyllensten, K. Helm-Bychowski, R.G. Higuchi, S.R. Palumbi, E.M. Prager, R.D. Sage, and M. Stoneking. 1985. Mitochondrial DNA in relation to evolutionary genetics. Biological Journal of the Linnaean Society 26: 375-400.


Stoneking, M., B. May, and J.E. Wright. 1981. Loss of duplicate gene expression in salmonids: evidence for a null allele polymorphism at the duplicate aspartate aminotransferase loci in brook trout (Salvelinus fontinalis). Biochemical Genetics 19: 1063-1077.

Stoneking, M., D.J. Wagner, and A.C. Hildebrand. 1981. Genetic evidence suggesting subspecific differences between northern and southern populations of brook trout (Salvelinus fontinalis). Copeia 1981: 810-819.


Wright, J.E., B. May, M. Stoneking, and G.M. Lee. 1980. Pseudolinkage of the duplicate loci for supernatant aspartate aminotransferase in brook trout, Salvelinus fontinalis. Journal of Heredity 71: 223-228.

May, B., M. Stoneking, and J.E. Wright. 1980. Joint segregation of biochemical loci in Salmonidae: II. Linkage associations from a hybridized Salvelinus genome (S. namaycush X S. fontinalis). Genetics 95: 707-726.


Stoneking, M., B. May, and J.E. Wright. 1979. Genetic variation, inheritance, and quaternary structure of malic enzyme in brook trout (Salvelinus fontinalis). Biochemical Genetics 17: 599-619.

May, B., J.E. Wright, and M. Stoneking. 1979. Joint segregation of biochemical loci in Salmonidae: results from experiments with Salvelinus and review of the literature on other species. Journal of the Fisheries Research Board of Canada 36: 1114-1128.

May, B., M. Stoneking, and J.E. Wright. 1979. Joint segregation of malate dehydrogenase and diaphorase loci in brown trout (Salmo trutta). Transactions of the American Fisheries Society 108: 373-377.


Vice Ombudsperson - Mark Stoneking

"Scientific honesty and the observance of the principle of good scientific practice are essential in all scientific work which seeks to expand our knowledge and which is intended to earn respect from the public." (Rules of Good Scientific Practice, MPS, 2009)

Ombudsperson: Shannon McPherron
Vice Ombudsperson: Mark Stoneking

As elected Ombudsperson and Vice Ombudsperson of the Max Planck Institute for Evolutionary Anthropology, we stand at your disposal in case you are experiencing or observing any kind of scientific misconduct, or if you need advice on the subject of good scientific practice.

Scientific misconduct includes: false statements, infringement of intellectual property, impairment of the research work of others, joint accountability. More information is provided in the following brochure:

Rules of Good Scientific Practice
March 20th, 2009
[pdf - English] [pdf - German]

Rules of Procedure in Case of Suspected Scientific Misconduct
November 24th, 2000
[pdf - English] [pdf - German]

Guidelines for the Deployment of Ombudspersons
February 16th, 2006
[pdf - English] [pdf - German]