Tracy Kivell

Pietrobelli, A., Marchi, D., Noerwidi, S., Alamsyah, N., Sutikna, T., Kivell, T. L., Skinner, M. M., & Tocheri, M. W. (2025). A new distal fibular fragment of Homo floresiensisand the first quantitative comparative analysis of proximal and distal fibular morphology in this species. Journal of Anatomy, 246(6), 869-891.
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Robbins, M. M., Drummond-Clarke, R. C., Robbins, A. M., Ostrofsky, K. R., & Kivell, T. L. (2025). Gorillas are arboreal apes. Current Biology, 35(12): e3, pp. 2974-2979.
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Syeda, S. M., Dunmore, C. J., Skinner, M. M., Berger, L. R., Churchill, S. E., Zipfel, B., & Kivell, T. L. (2025). Phalangeal cortical bone distribution reveals different dexterous and climbing behaviors in Australopithecus sediba and Homo naledi. Science Advances, 11(2): eadt1201.
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Tomkinson, G. R., Lang, J. J., Rubín, L., McGrath, R., Gower, B., Boyle, T., Klug, M. G., Mayhew, A. J., Blake, H. T., Ortega, F. B., Cadenas-Sanchez, C., Magnussen, C. G., Fraser, B. J., Kidokoro, T., Liu, Y., Christensen, K., Leong, D. P., Aadahl, M., Abdin, E., Alcazar, J., Alenazi, A., Alqahtani, B., Amaral, C. D. A., Amaral, T. L., Fernandes, A. A., Axelsson, P., Baldwin, J. N., Bammann, K., Barbosa, A. R., Bardo, A., Bimali, I., Bjerregaard, P., Bobak, M., Boreham, C. A., Bös, K., Marins, J. C. B., Burns, J., Capkova, N., Castillo-Martínez, L., Chen, L.-K., Choi, S. M., Choong, R. K., Confortin, S. C., Cooper, C., Correa-Bautista, J. E., Cournil, A., Cruz, G., de Bruin, E. D., De Paz, J. A., Moreira, B. D. S., Anjos, L. A. D., Reyna, M. C. E., Ferriolli, E., Forrester, G., Frolova, E., Gebre, A. K., Ghaleb, A. M., Gill, T. K., Gondo, Y., Gonzalez, M. C., Alvarez, C. G., Hannah, M. K., Harvey, N. C., Hogrel, J.-Y., Huemer, M.-T., Iidaka, T., Ingram, L. A., Jdanov, D. A., Keevil, V. L., Kemmler, W., Kenny, R. A., Kim, D.-Y., Kivell, T. L., Kjær, I. G., Kluttig, A., Kozakai, R., Langer, D., Larsen, L. A., Lee, W.-J., Leon, D. A., Lichtenstein, E., Little, B. B., Lourenço, R. A., Malhotra, R., Malina, R. M., Matsumoto, K., Mazor-Karsenty, T., McKay, M. J., McLoughlin, S., Mensegere, A. L., Mohammadian, M., Moreira, V. G., Murayama, H., Murray, A., Neri, A. L., Niessner, C., Othón, G. N., Olveira, G., Orchard, S. G., Pajak, A., Park, C. W., Pasco, J. A., Reyes, M. E. P., Pereira, L. S. M., Peters, A., Poon, E.-T.-C., Portela, M. C., Pratt, J., Ramírez-Vélez, R., Rodríguez-García, W., Ryan, J., San-Martín, M. A., Sánchez-Torralvo, F. J., Saremi, M., Schmidt-Trucksäss, A., Seino, S., Shah, S. A., Sim, M., Strand, B. H., Subramaniam, M., Suetta, C., Sui, S. X., Sundarakumar, J. S., Suzuki, K., Tamosiunas, A., Tan, M. P., Taniguchi, Y., Thorand, B., Turusheva, A., Tveter, A. T., Wagner, J., Wang, D., Warden, S. J., Wearing, J., Wee, S. L., Westbury, L. D., Wiśniowska-Szurlej, A., Woll, A., Yoshimura, N., & Yu, R. (2025). International norms for adult handgrip strength: A systematic review of data on 2.4 million adults aged 20 to 100+ years from 69 countries and regions. Journal of Sport and Health Science, 14: 101014.
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Bardo, A., D’Aout, K., Özçelebi, J., & Kivell, T. L. (2024). A helping hand: investigating 3D motion of human hand bones during Palaeolithic tool behaviours. Bulletins et mémoires de la Société d’Anthropologie de Paris BMSAP, 36(Suppl).
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Behringer, V., Sonnweber, R., Hohmann, G., Stevens, J. M. G., Verspeek, J., & Kivell, T. L. (2024). A non‐invasive measure of bone growth in mammals: Validating urinary CTX‐I as a bone resorption marker through long‐bone growth velocity in bonobos. Ecology and Evolution, 14: e70326.
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Cofran, Z., & Kivell, T. L. (2024). Adult Homo naledi hand skeleton points to life before birth. Comptes Rendus Palevol, 23(28), 441-451.
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Donati, G., Edginton, T., Bardo, A., Kivell, T. L., Ballieux, H., Stamate, C., & Forrester, G. S. (2024). Motor-sensory biases are associated with cognitive and social abilities in humans. Scientific Reports, 14: 14724.
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Drummond-Clarke, R. C., Kivell, T. L., Sarringhaus, L., Stewart, F. A., & Piel, A. K. (2024). Sex differences in positional behavior of chimpanzees (Pan troglodytes schweinfurthii) living in the dry and open habitat ofIssa Valley, Tanzania. American journal of biological anthropology, 185(3): e25007.
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Lukova, A., Dunmore, C. J., Bachmann, S., Synek, A., Pahr, D. H., Kivell, T. L., & Skinner, M. M. (2024). Trabecular architecture of the distal femur in extant hominids. Journal of Anatomy, 245(1), 156-180.
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Orr, C. M., Kivell, T. L., & Tocheri, M. W. (2024). Mary Marzke,PhD: A pioneer and innovator in the study of the evolution of the human hand. American Journal of Biological Anthropology, 183(13): e24892.
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Syeda, S. M., Tsegai, Z. J., Cazenave, M., Skinner, M. M., & Kivell, T. L. (2024). Cortical bone architecture of hominid intermediate phalanges reveals functional signals of locomotion and manipulation. American Journal of Biological Anthropology, 184(1): e24902.
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Tanner, S. B., Bardo, A., Davies, T. W., Dunmore, C. J., Johnston, R. E., Owen, N. J., Kivell, T. L., & Skinner, M. M. (2024). Variation and covariation of external shape and cross‐sectional geometry in the human metacarpus. American Journal of Biological Anthropology, 183(2): e24866.
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Bardo, A., Dunmore, C. J., Cornette, R., & Kivell, T. L. (2023). Morphological integration and shape covariation between the trapezium and first metacarpal among extant hominids. American Journal of Biological Anthropology.
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Cazenave, M., & Kivell, T. L. (2023). Challenges and perspectives on functional interpretations of australopith postcrania and the reconstruction of hominin locomotion. Journal of Human Evolution, 175: 103304.
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Dunmore, C. J., Bachmann, S., Synek, A., Pahr, D. H., Skinner, M. M., & Kivell, T. L. (2023). The deep trabecular structure of first metacarpals in extant hominids. American Journal of Biological Anthropology, 183(3): e24695.
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Syeda, S. M., Tsegai, Z. J., Cazenave, M., Skinner, M. M., & Kivell, T. L. (2023). Cortical bone distribution of the proximal phalanges in great apes: Implications for reconstructing manual behaviours. Journal of Anatomy, 243(5), 707-728.
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Bardo, A., Kivell, T. L., Donati, G., Ballieux, H., Stamate, C., Edginton, T., & Forrester, G. S. (2022). Recherche des interactions entre forme et fonction de la main chez les humains actuels dans la perspective de mieux apprécier l’évolution de la dextérité humaine = Research of the interactions in hand’s form and function in modern humans for a better understanding of the evolution of the human dexterity. Bulletins et mémoires de la société d'anthropologie de Paris, 34(S).
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Bardo, A., Town, K., Kivell, T. L., Donati, G., Ballieux, H., Stamate, C., Edginton, T., & Forrester, G. S. (2022). The precision of the human hand: Variability in pinch strength and manual dexterity. Symmetry, 14(1): 71.
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Bird, E. E., Kivell, T. L., & Skinner, M. M. (2022). Patterns of internal bone structure and functional adaptation in the hominoid scaphoid, lunate, and triquetrum. American Journal of Biological Anthropology, 177(2), 266-285.
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Cazenave, M., Kivell, T. L., Pina, M., Begun, D. R., & Skinner, M. M. (2022). Calcar femorale variation in extant and fossil hominids and its relationship with bipedalism. American Journal of Biological Anthropology, 177(S73), 29-29.
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Cazenave, M., Kivell, T. L., Pina, M., Begun, D. R., & Skinner, M. M. (2022). Calcar femorale variation in extant and fossil hominids: implications for identifying bipedal locomotion in fossil hominins. Journal of Human Evolution, 167: 103183.
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Deckers, K., Tsegai, Z. J., Skinner, M. M., Zeininger, A., & Kivell, T. L. (2022). Ontogenetic changes to metacarpal trabecular bone structure in mountain and western lowland gorillas. Journal of Anatomy, 241(1), 82-100.
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Drummond-Clarke, R. C., Kivell, T. L., Sarringhaus, L., Stewart, F. A., Humle, T., & Piel, A. K. (2022). Wild chimpanzee behavior suggests that a savanna-mosaic habitat did not support the emergence of hominin terrestrial bipedalism. Science Advances, 8: eadd9752.
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Kivell, T. L., Baraki, N., Lockwood, V., Williams‐Hatala, E. M., & Wood, B. A. (2022). Form, function and evolution of the human hand. American Journal of Biological Anthropology, 181(S76), 6-57.
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Lockwood, V. A., Lu, S.-C., Winter, S., & Kivell, T. L. (2022). Pressure generated by the modern human hand during vertical climbing. American Journal of Biological Anthropology, 177(S73), 108-109.
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Tsegai, Z. J., Gardner, P., Chamberlain, B., Desilva, J. M., Zipfel, B., Kivell, T. L., & Skinner, M. M. (2022). Locomotor variability in Sterkfontein Member 4: Analysis of the external shape and internal bone structure of the StW 562 and StW 59 first metatarsals. American Journal of Biological Anthropology, 177(S73), 185-185.
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Arias-Martorell, J., Dunmore, C. J., Skinner, M. M., & Kivell, T. L. (2021). The trabecular structure of the proximal ulna in hominoids. American Journal of Physical Anthropology, 174(S71), 5-5.
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Arias-Martorell, J., Zeininger, A., & Kivell, T. L. (2021). Trabecular structure of the elbow reveals divergence in knuckle-walking biomechanical strategies of African apes. Evolution, 75(11), 2959-2971.
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Bardo, A., Kivell, T. L., Town, K., Donati, G., Ballieux, H., Stamate, C., Edginton, T., & Forrester, G. S. (2021). Get a grip: Variation in human hand grip strength and implications for human evolution. Symmetry, 13: 1142.
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Bird, E. E., Kivell, T. L., & Skinner, M. M. (2021). Cortical and trabecular bone structure of the hominoid capitate. Journal of Anatomy, 239(2), 351-373.
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Bowland, L. A., Scott, J. E., Kivell, T. L., Patel, B. A., Tocheri, M. W., & Orr, C. M. (2021). Homo naledi pollical metacarpal shaft morphology is distinctive and intermediate between that of australopiths and other members of the genus Homo. Journal of Human Evolution, 158: 103048.
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Kivell, T. L. (2021). Human evolution: Thumbs up for efficiency. Current Biology, 31, R289-R291.
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Bardo, A., Moncel, M.-H., Dunmore, C. J., Kivell, T. L., Pouydebat, E., & Cornette, R. (2020). The implications of thumb movements for Neanderthal and modern human manipulation. Scientific Reports, 10: 19323.
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Dunmore, C. J., Bardo, A., Skinner, M. M., & Kivell, T. L. (2020). Trabecular variation in the first metacarpal and manipulation in hominids. American Journal of Physical Anthropology, 171(2), 219-241.
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Dunmore, C. J., Skinner, M. M., Bardo, A., Berger, L. R., Hublin, J.-J., Pahr, D. H., Rosas, A., Stephens, N. B., & Kivell, T. L. (2020). The position of Australopithecus sediba within fossil hominin hand use diversity. Nature Ecology & Evolution, 4, 911-918.
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Georgiou, L., Dunmore, C. J., Bardo, A., Buck, L. T., Hublin, J.-J., Pahr, D. H., Stratford, D., Synek, A., Kivell, T. L., & Skinner, M. M. (2020). Evidence for habitual climbing in a Pleistocene hominin in South Africa. Proceedings of the National Academy of Sciences of the United States of America, 117(15): 8423.
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Lockwood, V. A., Lu, S.-C., Winter, S., & Kivell, T. L. (2020). Support size and thumb position preferences in humans during suspension and vertical climbing: Implications for hominin hand evolution. American Journal of Physical Anthropology, 171(S69), 164-164.
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Skinner, M. M., Georgiou, L., Stratford, D., Dunmore, C. J., Bardo, A., Buck, L. T., Hublin, J.-J., Pahr, D. H., Synek, A., & Kivell, T. L. (2020). Reply to Haeusler et al.: Internal structure of the femur provides robust evidence for locomotor and taxonomic diversity at Sterkfontein. Proceedings of the National Academy of Sciences, 117, 28570-28571.
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Synek, A., Lu, S.-C., Nauwelaerts, S., Pahr, D. H., & Kivell, T. L. (2020). Metacarpophalangeal joint loads during bonobo locomotion: Model predictions versus proxies. Journal of The Royal Society Interface, 17(164): 20200032.
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Williams-Hatala, E. M., Hatala, K. G., Key, A., Dunmore, C. J., & Kivell, T. L. (2020). Kinetics of stone tool production in novice and expert toolmakers. American Journal of Physical Anthropology, 171(S69), 308-309.
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Williams‐Hatala, E. M., Hatala, K. G., Key, A., Dunmore, C. J., Kasper, M., Gordon, M., & Kivell, T. L. (2020). Kinetics of stone tool production among novice and expert tool makers. American Journal of Physical Anthropology, 174(4), 714-727.
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Deckers, K. P., Dunmore, C. J., Skinner, M. M., & Kivell, T. L. (2019). Trabecular ontogeny of African ape third metacarpals. American Journal of Physical Anthropology, 168(S68), 55-55.
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Deckers, K., Tsegai, Z. J., Zeininger, A., Skinner, M. M., & Kivell, T. L. (2019). Ontogenetic changes in third metacarpal trabecular bone in western lowland (Gorilla gorilla) and mountain gorillas (Gorilla beringei). In Proceedings of the European Society for the study of Human Evolution (PESHE) (8, pp. 47).
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Dunmore, C. C., Kivell, T. L., Stephens, N. B., Bardo, A., Hublin, J.-J., & Skinner, M. M. (2019). First metacarpal trabecular morphology: Implications for thumb use in great apes and Australopithecus. American Journal of Physical Anthropology, 168(S68), 64-64.
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Dunmore, C. J., Kivell, T. L., Bardo, A., & Skinner, M. (2019). Metacarpal trabecular bone varies with distinct hand-positions used in hominid locomotion. Journal of Anatomy, 235(1), 45-66.
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Galletta, L., Stephens, N. B., Bardo, A., Kivell, T. L., & Marchi, D. (2019). Three-dimensional geometric morphometric analysis of the first metacarpal distal articular surface in humans, great apes and fossil hominins. Journal of Human Evolution, 132, 119-136.
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Georgiou, L., Kivell, T. L., Pahr, D. H., Buck, L. T., & Skinner, M. M. (2019). Trabecular architecture of the great ape and human femoral head. Journal of Anatomy, 234(5), 679-693.
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Kivell, T. L. (2019). Fossil ape hints at how bipedal walking evolved. Nature, 575, 445-446.
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Kivell, T. L., Dunmore, C. J., Lu, S. C., Synek, A., Bardo, A., Bird, E. E., Decker, K. A. P., Key, A. J. M., Pahr, D. H., & Skinner, M. M. (2019). Signals of locomotion and manipulation in the internal trabecular bone structure of extant hominoids and fossil hominins. Journal of Morphology, 280(S1), S17-S18.
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Ríos, L., Kivell, T. L., Lalueza-Fox, C., Estalrrich, A., García-Tabernero, A., Huguet, R., Quintino, Y., de la Rasilla, M., & Rosas, A. (2019). Skeletal anomalies in the Neandertal family of El Sidrón (Spain) support a role of inbreeding in Neandertal extinction. Scientific Reports, 9: 1697.
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Synek, A., Dunmore, C. J., Kivell, T. L., Skinner, M. M., & Pahr, D. H. (2019). Inverse remodelling algorithm identifies habitual manual activities of primates based on metacarpal bone architecture. Biomechanics and Modeling in Mechanobiology, 18(2), 399-410.
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Synek, A., Lu, S.-C., Vereecke, E. E., Nauwelaerts, S., Kivell, T. L., & Pahr, D. H. (2019). Musculoskeletal models of a human and bonobo finger: parameter identification and comparison to in vitro experiments. PeerJ, 7: e7470.
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Tsegai, Z. J., Kivell, T. L., & Skinner, M. M. (2019). Trabecular structure of the third metatarsal head distinguishes between a grasping and non-grasping foot. In Proceedings of the European Society for the study of Human Evolution (PESHE) (8, pp. 188).
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Bardo, A., Vigouroux, L., Kivell, T. L., & Pouydebat, E. (2018). The impact of hand proportions on tool grip abilities in humans, great apes and fossil hominins: A biomechanical analysis using musculoskeletal simulation. Journal of Human Evolution, 125, 106-121.
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Bowland, L. A., Scott, J. E., Kivell, T. L., Patel, B. A., Tocheri, M. W., & Orr, C. M. (2018). Pollical metacarpal shaft morphology in Homo naledi: a 3D geometric morphometric analysis. American Journal of Physical Anthropology, 165(S66), 34-34.
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Dunmore, C. J., Kivell, T. L., Pahr, D. H., & Skinner, M. M. (2018). Trabecular architecture across the metacarpus reflects different locomotor strategies in hominoids. American Journal of Physical Anthropology, 165(S66), 73.
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Georgiou, L., Kivell, T. L., Pahr, D. H., & Skinner, M. M. (2018). Trabecular bone patterning in the hominoid distal femur. PeerJ, 6: e5156.
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Georgiou, L., Kivell, T. L., Pahr, D. H., & Skinner, M. M. (2018). Trabecular bone structure of the distal femur in great apes. American Journal of Physical Anthropology, 165(S66), 97-97.
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Groucutt, H. S., Grün, R., Zalmout, I. A. S., Drake, N. A., Armitage, S. J., Candy, I., Clark-Wilson, R., Louys, J., Breeze, P. S., Duval, M., Buck, L. T., Kivell, T. L., Pomeroy, E., Stephens, N. B., Stock, J. T., Stewart, M., Price, G. J., Kinsley, L., Sung, W. W., Alsharekh, A., Al-Omari, A., Zahir, M., Memesh, A. M., Abdulshakoor, A. J., Al-Masari, A. M., Bahameem, A. A., Murayyi, K. M. S. A., Zahrani, B., Scerri, E. L. M., & Petraglia, M. D. (2018). Homo sapiens in Arabia by 85,000 years ago. Nature Ecology & Evolution, 2, 800-809.
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Key, A., Merritt, S. R., & Kivell, T. L. (2018). Hand grip diversity and frequency during the use of Lower Palaeolithic stone cutting-tools. Journal of Human Evolution, 125, 137-158.
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Kivell, T. L., Davenport, R., Hublin, J.-J., Thackeray, J. F., & Skinner, M. M. (2018). Trabecular architecture and joint loading of the proximal humerus in extant hominoids, Ateles, and Australopithecus africanus. American Journal of Physical Anthropology, 167(2), 348-365.
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Kivell, T. L., Rosas, A., Estalrrich, A., Huguet, R., García-Tabernero, A., Ríos, L., & de la Rasilla, M. (2018). New Neandertal wrist bones from El Sidrón, Spain (1994–2009). Journal of Human Evolution, 114, 45-75.
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Komza, K., Kivell, T. L., & Skinner, M. M. (2018). Unique forms of locomotion in Swartkrans hominins: An analysis of the trabecular structure of the first metatarsal. American Journal of Physical Anthropology, 165(S66), 144-144.
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Lu, S. C., Vereecke, E. E., Synek, A., Pahr, D. H., & Kivell, T. L. (2018). A novel experimental design for the measurement of metacarpal bone loading and deformation and fingertip force. PeerJ, 5480.
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Neufuss, J., Robbins, M. M., Bäumer, J., Humle, T., & Kivell, T. L. (2018). Gait characteristics of vertical climbing in mountain gorillas and chimpanzees. Journal of Zoology, 306(2), 129-138.
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Neufuss, J., Robbins, M. M., Bäumer, J., Humle, T., & Kivell, T. L. (2018). Manual skills for food processing by mountain gorillas (Gorilla beringei beringei) in Bwindi Impenetrable National Park, Uganda. Biological Journal of the Linnean Society, 127(3), 543-562.
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Samuel, D. S., Nauwelaerts, S., Stevens, J. M. G., & Kivell, T. L. (2018). Hand pressures during arboreal locomotion in captive bonobos (Pan paniscus). Journal of Experimental Biology, 221(8): jeb170910.
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Stephens, N. B., Kivell, T. L., Pahr, D., Hublin, J.-J., & Skinner, M. M. (2018). Trabecular bone patterning across the human hand. Journal of Human Evolution, 123, 1-23.
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Tsegai, Z. J., Skinner, M. M., Pahr, D. H., Hublin, J.-J., & Kivell, T. L. (2018). Ontogeny and variability of trabecular bone in the chimpanzee humerus, femur and tibia. American Journal of Physical Anthropology, 167(4), 713-736.
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Tsegai, Z. J., Skinner, M. M., Pahr, D. H., Hublin, J.-J., & Kivell, T. L. (2018). Systemic patterns of trabecular bone across the human and chimpanzee skeleton. Journal of Anatomy, 232(4), 641-656.
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Tsegai, Z. J., Skinner, M. M., Pahr, D., Hublin, J.-J., & Kivell, T. L. (2018). Trabecular bone ontogeny in the forelimb and hindlimb of chimpanzees. In Proceedings of the European Society for the study of Human Evolution (PESHE) (7, pp. 190).
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Williams-Hatala, E. M., Hatala, K. G., Gordon, M., Key, A., Kasper, M., & Kivell, T. L. (2018). The manual pressures of stone tool behaviors and their implications for the evolution of the human hand. Journal of Human Evolution, 119, 14-26.
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Arias-Martorell, J., Davenport, R., Kivell, T. L., & Skinner, M. M. (2017). Trabecular architecture of the hominoid humerus. American Journal of Physical Anthropology, 162(S64), 103-103.
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Dunmore, C. J., Key, A. J., Williams-Hatala, E. M., & Kivell, T. L. (2017). Is there a relationship between flake form and manual pressure during stone tool production? An experimental test. In Proceedings of the European Society for the study of Human Evolution (PESHE) (5, pp. 84).
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Dunmore, C. J., Kivell, T. L., Pahr, D., & Skinner, M. M. (2017). Trabecular morphology across the hominoid metacarpus reflects distinct locomotor strategies. In Proceedings of the European Society for the study of Human Evolution (PESHE) (6, pp. 58).
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Hawks, J., Elliott, M., Schmid, P., Churchill, S. E., Ruiter, D. J. d., Roberts, E. M., Hilbert-Wolf, H., Garvin, H. M., Williams, S. A., Delezene, L. K., Feuerriegel, E. M., Randolph-Quinney, P., Kivell, T. L., Laird, M. F., Tawane, G., DeSilva, J. M., Bailey, S. E., Brophy, J. K., Meyer, M. R., Skinner, M. M., Tocheri, M. W., VanSickle, C., Walker, C. S., Campbell, T. L., Kuhn, B., Kruger, A., Tucker, S., Gurtov, A., Hlophe, N., Hunter, R., Morris, H., Peixotto, B., Ramalepa, M., Rooyen, D. v., Tsikoane, M., Boshoff, P., Dirks, P. H., & Berger, L. R. (2017). New fossil remains of Homo naledi from the Lesedi Chamber, South Africa. eLife, 6: e24232.
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Kivell, T. L., Estalrrich, A., Huguet, R., Garcia-Tabernero, A., Rios, L., de la Rasilla, M., & Rosas, A. (2017). It's all in the wrist: New Neandertal carpal bones from El Sidrón (Asturias, Spain). American Journal of Physical Anthropology, 162(S64), 246-246.
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Neufuss, J., Humle, T., Cremaschi, A., & Kivell, T. L. (2017). Nut-cracking behaviour in wild-born, rehabilitated bonobos (Pan paniscus): A comprehensive study of hand-preference, hand grips and efficiency. American Journal of Primatology, 79(2), 1-16.
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Neufuss, J., Robbins, M. M., Baeumer, J., Humle, T., & Kivell, T. L. (2017). Comparison of hand use and forelimb posture during vertical climbing in mountain gorillas (Gorilla beringei beringei) and chimpanzees (Pan troglodytes). American Journal of Physical Anthropology, 164(4), 651-664.
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Neufuss, J., Robbins, M. M., Bäumer, J., Humle, T., & Kivell, T. L. (2017). Comparison of hand use and forelimb mechanics of vertical climbing in wild mountain gorillas and free-ranging chimpanzees. Folia primatologica, 88(2), 138-139.
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Stephens, N. B., Kivell, T. L., Pahr, D., Hublin, J.-J., & Skinner, M. M. (2017). Trabecular bone patterning across the human hand: Implications for reconstructing behavior and manipulation in past populations. In Proceedings of the European Society for the study of Human Evolution (PESHE) (6, pp. 190).
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Tsegai, Z. J., Skinner, M. M., Gee, A. H., Pahr, D. H., Treece, G. M., Hublin, J.-J., & Kivell, T. L. (2017). Trabecular and cortical bone structure of the talus and distal tibia in Pan and Homo. American Journal of Physical Anthropology, 163(4), 784-805.
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Tsegai, Z. J., Stephens, N. B., Treece, G. M., Skinner, M. M., Kivell, T. L., & Gee, A. H. (2017). Cortical bone mapping: An application to hand and foot bones in hominoids. Comptes Rendus Palevol, 16(5-6), 690-701.
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Williams-Hatala, E. M., Hatala, K. G., Gordon, M., Kasper, M., & Kivell, T. L. (2017). The biochemanics of stone tool behaviors and imlications for the evolution of the human hand. American Journal of Physical Anthropology, 162(S64), 411-411.
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Williams-Hatala, E. M., Hatala, K. G., Gordon, M., Kasper, M., & Kivell, T. L. (2017). The biomechanics of stone tool behaviors and implications for the evolution of the human hand. American Journal of Physical Anthropology, 162(S64), 411-411.
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Behringer, V., Stevens, J. M. G., Kivell, T. L., Neufuss, J., Boesch, C., & Hohmann, G. (2016). Within arm's reach: Measuring forearm length to assess growth patterns in captive bonobos and chimpanzees. American Journal of Physical Anthropology, 161(1), 37-43.
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Kivell, T. L. (2016). A review of trabecular bone functional adaptation: What have we learned from trabecular analyses in extant hominoids and what can we apply to fossils? Journal of Anatomy, 228(4), 569-594.
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Kivell, T. L. (2016). The primate wrist. In T. L. Kivell, P. Lemelin, B. G. Richmond, & D. Schmitt (Eds.), The evolution of the primate hand: Anatomical, developmental, functional, and paleontological evidence (pp. 17-54). New York [u.a.]: Springer.
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Kivell, T. L., Deane, A. S., Tocheri, M. W., Orr, C. M., Schmid, P., Hawks, J., Berger, L. R., & Churchill, S. E. (2016). Functional interpretation of the Homo naledi hand. American Journal of Physical Anthropology, 159(S62), 192-192.
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Kivell, T. L., Lemelin, P., Richmond, B. G., & Schmitt, D. (Eds.). (2016). The evolution of the primate hand: Anatomical, developmental, functional, and paleontological evidence. New York [u.a.]: Springer.
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Kivell, T. L., Lemelin, P., Richmond, B. G., & Schmitt, D. (2016). Introduction. In T. L. Kivell, P. Lemelin, B. G. Richmond, & D. Schmitt (Eds.), The evolution of the primate hand: Anatomical, developmental, functional, and paleontological evidence (pp. 1-3). New York [u.a.]: Springer.
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Schmitt, D., Zeininger, A., Hamrick, E., Snyder, M. L., Kivell, T. L., & Wunderlich, R. E. (2016). Gorilla limb kinematics and hominoid locomotor diversity: Implications for hominin locomotor evolution. American Journal of Physical Anthropology, 159(S62), 282-282.
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Schoonaert, K., D'Août, K., Samuel, D., Talloen, W., Nauwelaerts, S., Kivell, T. L., & Aerts, P. (2016). Gait characteristics and spatio-temporal variables of climbing in bonobos (Pan paniscus). American Journal of Primatology, 78(11), 1165-1177.
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Stephens, N. B., Kivell, T. L., Gross, T., Pahr, D. H., Lazenby, R. A., Hublin, J.-J., Hershkovitz, I., & Skinner, M. M. (2016). Trabecular architecture in the thumb of Pan and Homo: Implications for investigating hand use, loading, and hand preference in the fossil record. American Journal of Physical Anthropology, 161(4), 603-619.
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Stephens, N. B., Kivell, T. L., Pahr, D. H., Gee, A. H., Treece, G. M., Hublin, J.-J., & Skinner, M. M. (2016). Signals of loading and function in the human hand: A multi-method analysis of the external cortical and internal trabecular bone of the metacarpals. American Journal of Physical Anthropology, 159(S62), 302-303.
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Tsegai, Z. J., Skinner, M. M., Pahr, D. H., Hublin, J.-J., & Kivell, T. L. (2016). Systemic patterns of trabecular structure in Homo and Pan: Evaluating inter- and intraspecific variability across anatomical sites. American Journal of Physical Anthropology, 159(S62), 318-319.
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Williams-Hatala, E. M., Key, A., Stephens, N. B., & Kivell, T. L. (2016). Predictions for an osteological signature of stone tool behaviors in hard tissue anatomy. American Journal of Physical Anthropology, 159(S62), 335-335.
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Berger, L. R., Hawks, J., de Ruiter, D. J., Churchill, S. E., Schmid, P., Delezene, L. K., Kivell, T. L., Garvin, H. M., Williams, S. A., DeSilva, J. M., Skinner, M. M., Musiba, C. M., Cameron, N., Holliday, T. W., Harcourt-Smith, W., Ackermann, R. R., Bastir, M., Bogin, B., Bolter, D., Brophy, J., Cofran, Z. D., Congdon, K. A., Deane, A. S., Dembo, M., Drapeau, M., Elliott, M. C., Feuerriegel, E. M., Garcia-Martinez, D., Green, D. J., Gurtov, A., Irish, J. D., Kruger, A., Laird, M. F., Marchi, D., Meyer, M. R., Nalla, S., Negash, E. W., Orr, C. M., Radovcic, D., Schroeder, L., Scott, J. E., Throckmorton, Z., Tocheri, M. W., VanSickle, C., Walker, C. S., Wei, P., & Zipfel, B. (2015). Homo naledi, a new species of the genus Homo from the Dinaledi Chamber, South Africa. eLife, e09560.
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Chirchir, H., Kivell, T. L., Ruff, C. B., Hublin, J.-J., Carlson, K. J., Zipfel, B., & Richmond, B. G. (2015). Recent origin of low trabecular bone density in modern humans. Proceedings of the National Academy of Sciences of the United States of America, 112(2), 366-371.
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Feix, T., Kivell, T. L., Pouydebat, E., & Dollar, A. M. (2015). Estimating thumb–index finger precision grip and manipulation potential in extant and fossil primates. Interface: Journal of the Royal Society, 12(106): 0176.
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Kivell, T. L. (2015). Evidence in hand: Recent discoveries and the early evolution of human manual manipulation. Philosophical Transactions of the Royal Society of London, Series B: Biological Sciences, 370(1682): 20150105.
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Kivell, T. L., Deane, A. S., Tocheri, M. W., Orr, C. M., Schmid, P., Hawks, J., Berger, L. R., & Churchill, S. E. (2015). The hand of Homo naledi. Nature Communications, 6: 8431.
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Miszkiewicz, J. J., Kivell, T. L., Schlecht, S. H., & Mahoney, P. (2015). Investigating the extent to which entheseal changes reflect bone remodeling at the modern human femoral midshaft. American Journal of Physical Anthropology, 156(S60), 227-227.
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Neufuss, J., Humle, T., Deschner, T., Robbins, M. M., Sirianni, G., Boesch, C., & Kivell, T. L. (2015). Diversity of hand grips and laterality in wild African apes. Folia primatologica, 86(4), 329.
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Skinner, M. M., Stephens, N. B., Tsegai, Z. J., Foote, A. C., Nguyen, H. N., Gross, T., Pahr, D. H., Hublin, J.-J., & Kivell, T. L. (2015). Human-like hand use in Australopithecus africanus. Science, 347(6220), 395-399.
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Skinner, M. M., Stephens, N. B., Tsegai, Z. J., Foote, A. C., Nguyen, H. N., Gross, T., Pahr, D. H., Hublin, J.-J., & Kivell, T. L. (2015). Response to Comment on “Human-like hand use in Australopithecus africanus”. Science Magazine, 348(6239), 1101 -1101.
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Tsegai, Z. J., Skinner, M. M., Pahr, D. H., Hublin, J.-J., & Kivell, T. L. (2015). Trabecular bone architecture and distribution in the talus and distal tibia of Homo and Pan. In 5th Annual Meeting of the European Society for the Study of Human Evolution (ESHE) (pp. 221-221). European Society for the Study of Human Evolution (ESHE).
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Gross, T., Kivell, T. L., Skinner, M. M., Nguyen, H. N., & Pahr, D. H. (2014). A CT-image-based framework for the holistic analysis of cortical and trabecular bone morphology. Palaeontologia Electronica, 17(3): 17.3.33A. Retrieved from http://palaeo-electronica.org/content/2014/889-holistic-analysis-of-bone.
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Kivell, T. L., Nguyen, H. N., Tsegai, Z. J., Gross, T., Pahr, D. H., Hublin, J.-J., & Skinner, M. M. (2014). Fossils, trabeculae and finite element modelling: A holistic approach to reconstructing behaviour in the past. American Journal of Physical Anthropology, 153(S58), 158.
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Nguyen, H. N., Pahr, D. H., Gross, T., Skinner, M. M., & Kivell, T. L. (2014). Micro-finite element (μFE) modeling of the siamang (Symphalangus syndactylus) third proximal phalanx: The functional role of curvature and the flexor sheath ridge. Journal of Human Evolution, 67, 60-75.
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Pouydebat, E., Fragaszy, D. M., & Kivell, T. L. (2014). Grasping in primates: For feeding, moving and human specificities = Saisir chez les primates: Se nourrir, se deplacer et les specificités humaines. Bulletins et mémoires de la Société d'Anthropologie de Paris; BMSAP, 26(3-4), 129-133.
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Schilling, A.-M., Tofanelli, S., Hublin, J.-J., & Kivell, T. L. (2014). Trabecular bone structure in the primate wrist. Journal of Morphology, 275(5), 572-585.
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Kivell, T. L., Barros, A. P., & Smaers, J. B. (2013). A phylogenetically-integrated morphological analysis of the hominoid wrist. American Journal of Physical Anthropology, Suppl. 56, Special Issue, 168.
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Kivell, T. L., Barros, A. P., & Smaers, J. B. (2013). Different evolutionary pathways underlie the morphology of wrist bones in hominoids. BMC Evolutionary Biology, 13: 229.
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Kivell, T. L., Guimont, I., & Wall, C. E. (2013). Sex-Related Shape Dimorphism in the Human Radiocarpal and Midcarpal Joints. The Anatomical Record, 226(1), 19-30.
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Neufuss, J., Deschner, T., Robbins, M., Pouydebat, E., & Kivell, T. (2013). Hand use during non-locomotor behaviours in the wild African Apes. Folia primatologica, 84, 307.
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Nguyen, H. N., Pahr, D. H., Gross, T., Skinner, M. M., & Kivell, T. L. (2013). The biomechanical role of trabecular bone in the siamang (Symphalangus syndactylus) manual proximal phalanx. In European Society for the study of Human Evolution (Ed.), Proceedings of the European Society for the study of Human Evolution 2 (PESHE 2) (pp. 160-160). Retrieved from http://eshe.eu/static/eshe/files/ESHE_Vienna_2013_Abstracts.pdf.
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Pouydebat, E., Reghem, E., & Kivell, T. L. (2013). Eco-ethological and kinematical approaches to grasping in primates. Folia primatologica, 84(3-5), 314-314.
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Skinner, M. M., Kivell, T. L., Potze, S., & Hublin, J.-J. (2013). Microtomographic archive of fossil hominin specimens from Kromdraai B, South Africa. Journal of Human Evolution, 64(5), 434-447.
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Stephens, N. B., Kivell, T. L., Nguyen, H. N., Pahr, D. H., Gross, T., Hublin, J.-J., & Skinner, M. M. (2013). Trabecular bone architecture in the thumb of recent Homo sapiens, Pan, and Late Pleistocene Homo: taxonomic differences and evidence for handedness. American Journal of Physical Anthropology, 150(S56), 262-263.
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Stephens, N., Kivell, T. L., Nguyen, H. N., Gross, T., Pahr, D. H., Hublin, J.-J., & Skinner, M. M. (2013). Trabecular bone architecture in the thumb of Homo and Pan: Handedness and evidence for taxonomic differences. In European Society for the study of Human Evolution (Ed.), Proceedings of the European Society for the study of Human Evolution 2 (PESHE 2) (pp. 215-215). Retrieved from http://eshe.eu/static/eshe/files/ESHE_Vienna_2013_Abstracts.pdf.
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Tsegai, Z. J., Kivell, T. L., Gross, T., Nguyen, H. N., Pahr, D. H., & Skinner, M. M. (2013). Identifying behavioural signals in trabecular bone structure of the hominoid third metacarpal. Folia primatologica, 84, 334.
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Tsegai, Z. J., Kivell, T. L., Gross, T., Nguyen, H. N., Pahr, D. H., & Skinner, M. M. (2013). Trabecular bone distribution in the hominoid third metacarpal head reflects predicted loaded joint position. In European Society for the study of Human Evolution (Ed.), Proceedings of the European Society for the study of Human Evolution 2 (PESHE 2) (pp. 227-227). Retrieved from http://eshe.eu/static/eshe/files/ESHE_Vienna_2013_Abstracts.pdf.
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Tsegai, Z. J., Kivell, T. L., Gross, T., Nguyen, H. N., Pahr, D. H., Smaers, J. B., & Skinner, M. M. (2013). Trabecular bone structure correlates with hand posture and use in hominoids. PLoS One, 8(11): e78781.
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Kivell, T. L., Skinner, M. M., Lazenby, R. L., & Hublin, J.-J. (2012). Trabecular architecture of fossil hominin first metacarpals. American Journal of Physical Anthropology, 147(S54), 182-182.
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Nguyen, H. N., Kivell, T. L., Skinner, M. M., Pahr, D. H., & Hublin, J.-J. (2012). Three-dimensional microstructure finite element modelling of the third proximal phalanx in the siamang (Symphalangus syndactylus). Poster presented at 2nd Annual Meeting of the European Society for the Study of Human Evolution, Bordeaux.
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Begun, D. R., & Kivell, T. L. (2011). Knuckle-walking in Sivapithecus? The combined effects of homology and homoplasy with possible implications for pongine dispersals. Journal of Human Evolution, 60(2), 158-170.
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Berger, L., Kibii, J., Churchill, S. E., Schmid, P., Carlson, K., deKlerk, B., deRuiter, D., Holliday, T., Kivell, T. L., Gurche, J., Zipfel, B., DaSilva, J., & Kidd, R. (2011). New remains of Australopithecus sediba from the Malapa site, South Africa. American Journal of Physical Anthropology, 144(S52), 88-88.
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Chino, J. P., Lee, W. R., Madden, R., Sims, E. L., Kivell, T. L., Doyle, S. K., Mitchell, T. L., Hoppenworth, E. J., & Marks, L. B. (2011). Teaching the Anatomy of Oncology: Evaluating the Impact of a Dedicated Oncoanatomy Course. International Journal of Radiation: Oncology, Biology, Physics, 79(3), 853-859.
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Grote, S., Call, J., & Kivell, T. L. (2011). Who is more bipedal? Positional behaviour in captive bonobos and chimpanzees.
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Kivell, T. L. (2011). A comparative analysis of the hominin triquetrum (SKX 3498) from Swartkrans, South Africa. South African Journal of Science, 107(5/6): 515, pp. 1-10.
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Kivell, T. L., Kibii, J. M., Churchill, S. E., & Berger, L. R. (2011). Evolution of the hominin hand: old and new evidence from the Plio-Pleistocene.
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Kivell, T. L., Kibii, J. M., Churchill, S. E., Schmid, P., & Berger, L. R. (2011). Australopithecus sediba Hand Demonstrates Mosaic Evolution of Locomotor and Manipulative Abilities. Science, 333(6048), 1411-1417.
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Kivell, T. L., Skinner, M. M., Lazenby, R., & Hublin, J.-J. (2011). Methodological considerations for analyzing trabecular architecture: an example from the primate hand. Journal of Anatomy, 218(2), 209-225.
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Kivell, T. L., Skinner, M. M., Lazenby, R., & Hublin, J.-J. (2011). Trabecular architecture of the hominoid carpus. The FASEB Journal, 25(Supplement): 183.7.
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Kivell, T. L., Skinner, M. M., Lazenby, R., & Hublin, J.-J. (2011). Trabecular bone structure of the fossil hominin first metacarpals. In Abstracts of the 1st Annual Meeting of the European Society for the Study of Human Evolution.
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Lazenby, R. A., Skinner, M. M., Kivell, T. L., & Hublin, J.-J. (2011). Scaling VOI Size in 3D lCT Studies of Trabecular Bone: A Test of the Over-Sampling Hypothesis. American Journal of Physical Anthropology, 144(2), 196-203.
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Auerbach, B. M., Case, D. T., Kivell, T. L., & Garofalo, E. M. (2010). Appendicular Asymmetry Altogether: Gradients of Directional Asymmetry through the Human Upper Limb [Abstract].
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Kivell, T. L., & Guimont, I. (2010). A comparative analysis of sexual shape dimorphism in the human midcarpal joint. American Journal of Physical Anthropology, 141(S50), 144-144.
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Kivell, T. L., Schmitt, D., & Wunderlich, R. E. (2010). Hand and foot pressures in the aye-aye (Daubentonia madagascariensis) reveal novel biomechanical trade-offs required for walking on gracile digits. Journal of Experimental Biology, 213(9), 1549-1557.
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Parker, L. R., Wunderlich, R. E., & Kivell, T. L. (2010). Manual and pedal pressures during sloped quadrupedal locomotion in Lemur catta and Varecia rubra. American Journal of Physical Anthropology, 141(S50), 185-185.
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Kivell, T. L., & Begun, D. R. (2009). New primate carpal bones from Rudabánya (late Miocene, Hungary): Taxonomic and functional implications. Journal of Human Evolution, 57(6), 697-709.
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Kivell, T. L., & Guimont, I. (2009). Evolutionary implications of sexual dimorphism in the human midcarpal joint. Talk presented at Annual Meeting at Experimental Biology of the American Association of Anatomists. New Orleans. 2009-04 - 2009-04.
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Kivell, T. L., & Schmitt, D. (2009). Independent evolution of knuckle-walking in African apes shows that humans did not evolve from a knuckle-walking ancestor. PNAS, 106(34), 14241-14246.
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Kivell, T. L., Doyle, S. K., Madden, R. H., Mitchell, T. L., & Sims, E. L. (2009). An interactive method for teaching anatomy of the human eye for ophthalmology rotations using suid eyes. Talk presented at Annual Meeting at Experimental Biology of the American Association of Anatomists. New Orleans. 2009-04 - 2009-04.
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Kivell, T. L., Doyle, S. K., Madden, R. H., Mitchell, T. L., & Sims, E. L. (2009). An interactive method for teaching anatomy of the human eye for medical students in ophthalmology clinical rotations. Anatomical Sciences Education, 2(4), 173-178.
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Kivell, T. L., Kramer, E. M., & Wunderlich, R. E. (2009). Aye-aye hand posture and loading of their specialized digits during quadrupedal locomotion. Talk presented at The 31st Annual Meeting of the Society of Integrative and Comparative Biology (SICB). Boston, Massachusetts. 2009-01-03 - 2009-01-07.
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Kivell, T. L., Wunderlich, R. E., & Kramer, E. M. (2009). Aye-aye hand and foot postures and loading during quadrupedal locomotion. American Journal of Physical Anthropology, 138(S48), 166-166.
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Wunderlich, R. E., & Kivell, T. L. (2009). Postcranial ontogeny in captive sifaka (Propithecus verreauxi). American Journal of Physical Anthropology, 138(S48), 277-277.
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Kivell, T. L. (2008). Ontogeny of the hominoid carpus: Did we evolve from a knuckle-walking ancestor? American Journal of Physical Anthropology, 135(S46), 131-132.
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Kivell, T. L., Kramer, E. M., & Wunderlich, R. E. (2008). Aye-aye (Daubentonia madagascariensis) hand posture and loading of their specialized digits during quadrupedal locomotion. Talk presented at Southeast regional SICB meeting for the Divisions of Vertebrate Morphology and Comparative Biomechanics. Clemson University, South Carolina. 2008-10 - 2008-10.
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Guimont, I., & Kivell, T. L. (2007). Inter- and intraspecific differences in primate carpal morphology. American Journal of Physical Anthropology, 132(S44), 121-121.
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Kivell, T. L. (2007). Discovering the past: Sibudu Cave. In M. Chazan (Ed.), World prehistory and archaeology: Pathways through time (pp. 142-143). Boston: Allyn & Bacon.
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Kivell, T. L. (2007). Ontogeny of the hominoid midcarpal joint and implications for the origin of hominin bipedalism. PhD Thesis, University of Toronto, Toronto.
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Kivell, T. L., & Begun, D. R. (2007). Frequency and timing of scaphoid-centrale fusion in hominoids. Journal of Human Evolution, 52(3), 321-340.
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McGoogan, K., Kivell, T., Hutchison, M., Young, H., Blanchard, S., Keeth, M., & Lehman, S. M. (2007). Phylogenetic diversity and the conservation biogeography of African primates. Journal of Biogeography, 34(11), 1962-1974.
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Begun, D. R., Ward, C. V., Deane, A. C., Kivell, T. L., Nargolwalla, M. C., & Taylor, N. (2006). Stem hominine or hominid? The phylogeny and functional anatomy of Pierolapithecus catalaunicus. American Journal of Physical Anthropology, 129(S42), 63-63.
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Kivell, T. L., & Begun, D. R. (2006). Functional morphology of new Dryopithecus carpals from Rudabánya, Hungary. American Journal of Physical Anthropology, 129(S42), 114-114.
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Kivell, T. L. (2005). Phylogenetic and functional analysis of primate carpal ossification sequences: A test of two methods. American Journal of Physical Anthropology, 126(S40), 130-131.
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Begun, D. R., Kivell, T. L., & Kordos, L. (2003). New Miocene primate postcranial fossils from Rudabánya, Hungary. American Journal of Physical Anthropology, 120(S36), 63-64.
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Kivell, T. L. (2002). Reevaluation of Oreopithecus positional repertoire. Talk presented at The 30th Annual Meeting of the Canadian Association of Physical Anthropology. Ottawa, ON. 2002-10-24 - 2002-10-26.
BibTeX   Endnote   

Kivell, T. L. (2002). No longer enigmatic: Reevalutation of Oreopithecus bambolii positional repertoire. Master Thesis, University of Toronto, Toronto.
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